Time devised by von Buddenbrock [1] for animals that can detect vibration without a resonant organ (i.e. those without echte Gehörsinn).

This distinction is criticised by Pumphrey [2] : "This invention has had unfortunate consequences, because of the tendency to regard sensitivity to vibrations as a special kind of touch sense in quite a separate category from true hearing instead of regarding it as including true hearing, which would be both logical and consistent with what is known of the evolution of hearing organs."


  1. von Buddenbrock WD. Grundiss der vergleichenden Physiologie. Berlin: Borntraeger; 1937.
  2. Pumphrey RJ. Hearing in Insects. Biological Reviews. 1940;15(1):107 - 132. Available at:

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Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith