@article {58380, title = {Bioacoustics of poorly known Poecilimon taxa (Insecta: Orthoptera: Tettigoniidae) with redescriptions of P. pechevi and P. stschelkanovzevi}, journal = {Zootaxa}, volume = {4890}, year = {2020}, month = {Apr-12-2020}, pages = {535 - 553}, abstract = {

The genus Poecilimon has been object of several studies dealing with its systematics, sound communication and evolution. Yet, published data contradict in the classification, while many taxa are still insufficiently known. In the present study we supplement the knowledge of 13 poorly known species and one additional subspecies with data on their sound communication and/or morphology. Most species concerned here are classified within two acoustic groups. First group includes P. celebi, P. obtusicercus and P. toros, showing typical characteristics of the P. syriacus group. Second group is more heterogeneous. Among that, P. pechevi, P. armeniacus, P. harveyi, P. guichardi, P. haydari, P. doga, P. davisi and P. excisus present a song pattern similar to that of P. ampliatus. In addition, P. ataturki is also quite similar in basic song structure to the latter but differs in its fine song structure. Poecilimon stschelkanovzevi, with unclear relationships, is described morphologically in detail. The present study provides hints for solving some disagreement between recent revisions, proposing unification of the Poecilimon celebi with P. syriacus group and opening discussion based on close relationships of the groups ampliatus, armeniacus, davisi, luschani and the species P. guichardi on one hand, while the complex P. ataturki/glandifer shows significant differences on the other. The acoustic peculiarities of the ampliatus group allies are discussed in an evolutionary light.

}, keywords = {Barbitistini, bush-cricket, morphology, sound communication, sound production}, issn = {1175-5326}, doi = {10.11646/zootaxa.4890.410.11646/zootaxa.4890.4.6}, url = {https://www.biotaxa.org/Zootaxa/issue/view/zootaxa.4890.4https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4890.4.6https://www.biotaxa.org/Zootaxa/article/viewFile/65904/64346}, author = {Dragan Petrov Chobanov and Sevgili, Hasan and Klaus-Gerhard Heller} } @article {58028, title = {Poecilimon gerlindae spec. nov.{\textendash}a new bushcricket species of the Poecilimon propinquus-group (Orthoptera: Phaneropteridae) from Greece}, journal = {Articulata}, volume = {21}, year = {2006}, pages = {109-119}, chapter = {109}, abstract = {

Poecilimon gerlindae spec. nov. from Greek mainland is described. The new species is the eight known member of the P. propinquus-group (sensu Lehmann AW 1998). It is distinct with respect to male cerci, the corresponding female basal fold, body colouration, number of teeth on the stridulatory file and the num- ber of pulses in the song. Otherwise it resembles P. propinquus in body size and general song pattern.

}, author = {Lehmann, Arne W and Willemse, Fer and Klaus-Gerhard Heller} } @article {57963, title = {Review of Poecilimon species with inflated pronotum: description of four new taxa within an acoustically diverse group}, year = {2018}, abstract = {

Poecilimon Fischer (Orthoptera, Tettigoniidae) consists of about 130 species and a functional taxonomy of the genus re- quires arranging species into phylogenetic groups. However, this is a task that requires detailed empirical testing. This paper documents data on Poecilimon species with an inflated pronotum. The Poecilimon (Poecilimon) inflatus group is a lineage ranging along a narrow zone in the south-west corner of Anatolia and in Crete plus some other Aegean islands. By producing data of morphology and calling songs of males and responding songs of females the composition and intra- group diversity of P. inflatus group are studied. Both morphology and male calling and female responding songs, exhibited a considerable intra-group diversity. In the light of data obtained the following nomenclatural changes were made. The genus Parapoecilimon Karaba{\u g}, 1975 syn. n. is synonymised with Poecilimon Fischer and Parapoecilimon antalyaensis Karaba{\u g}, 1975 is given a new combination, Poecilimon antalyaensis (Karaba{\u g}, 1975) comb. n. Four new taxa are de- scribed in the group: (i) Poecilimon isopterus sp.n.; (ii) Poecilimon inflatus lyciae subsp. n.; (iii) Poecilimon antalyaensis myrae subsp. n., and (iv) Poecilimon antalyaensis anemurium subsp. n. The Poecilimon (P.) inflatus species group con- sists of the following species: (i) Poecilimon inflatus Brunner von Wattenwyl, 1891; (ii) Poecilimon martinae Heller, 2004; (iii) Poecilimon cretensis Werner, 1903; (iv) Poecilimon antalyaensis (Karaba{\u g}, 1975) comb. n.; (v) Poecilimon bilgeri Karaba{\u g}, 1953 and (vi) Poecilimon isopterus sp.n. The species group is defined by a combination of characters including the pronotum inflated in metazona and a male cerci mostly black in their apical 1/2\–1/3. The male calling song has typically two types of short syllables and the females respond acoustically with very short latency times. By using combination of morphology and song data we suggest three main lineages in the group: (i) P. bilgeri, (ii) P. antalyaensis and (iii) the other four species.

}, keywords = {Anatolia, Crete, Phaneropterinae, Poecilimon, Poecilimon inflatus group, Poecilimon isopterus sp. n.}, doi = {10.11646/zootaxa.4462.410.11646/zootaxa.4462.4.1}, url = {https://biotaxa.org/Zootaxa/issue/view/zootaxa.4462.4https://biotaxa.org/Zootaxa/article/view/zootaxa.4462.4.1}, author = {Kaya, Sarp and Dragan Petrov Chobanov and Klaus-Gerhard Heller and {\"O}zg{\"u}l Yahyao{\u g}lu} } @article {57942, title = {Morphology, bioacoustics and phylogeography of the Isophya major group (Orthoptera: Tettigoniidae: Phaneropterinae): A species complex occurring in Anatolia and Cyprus}, journal = {European Journal of Entomology}, volume = {103}, year = {2006}, pages = {657}, abstract = {

The species of the Isophya major-group (Orthoptera: Tettigoniidae: Phaneropterinae), are reviewed. I. major Brunner von Wattenwyl and I. mavromoustakisi Uvarov are redescribed. Two new species are described; Isophya mersinensis Sevgili \& \Çiplak sp. n. and Isophya salmani Sevgili \& Heller sp. n. from south Anatolia. Illustrations of morphology and male calling song are provided and a detailed comparison of the four species in the group is presented. We conclude that these four species constitute a natural group in sharing male cerci with two or more denticles not ordered in a line, a character unique to this species group within the genus. From an evaluation of their morphology and song characteristics, relationships among the species in the group are I. major + (I. mersinensis sp. n. + (I. salmani + I. mavromoustakisi). Based on the the distribution pattern and habitat preference, we concluded that the division of I. salmani and I. mavromoustakisi was a vicariant event resulting from the separation of Cyprus and Anatolia due to reflooding of the Mediterranean after the Messinian salinity crisis at the beginning of the Pliocene, around 5 My ago.

}, keywords = {Anatolia, Biogeography, Cyprus, Eastern Mediterranean, I. major, I. mavromoustakisi, I. mersinensis sp. n., I. salmani sp. n., Isophya, Orthoptera, Phaneropterinae, Turkey}, author = {Hasan Sevgili and {\c C}iplak, Battal and Klaus-Gerhard Heller and Demirsoy, Ali} } @article {53094, title = {Review of Poecilimon species with inflated pronotum: description of four new taxa within an acoustically diverse group}, journal = {Zootaxa}, volume = {4462}, year = {2018}, month = {May-08-2020}, pages = {451}, issn = {1175-5326}, doi = {10.11646/zootaxa.4462.4.1}, url = {https://biotaxa.org/Zootaxa/article/view/zootaxa.4462.4.1}, author = {Kaya, Sarp and Dragan Petrov Chobanov and Klaus-Gerhard Heller and {\"O}zg{\"u}l Yahyao{\u g}lu} } @article {52947, title = {Paraplangia sinespeculo, a new genus and species of bush-cricket, with notes on its biology and a key to the genera of Phaneropterinae (Orthoptera: Tettigonioidea) from Madagascar}, journal = {Journal of Orthoptera Research}, volume = {27}, year = {2018}, month = {Nov-07-2019}, pages = {143 - 153}, abstract = {

Madagascar is a well-known hotspot of biodiversity. However, many Orthoptera, and especially the Tettigonioidea, belong to little-studied groups. Here we describe a new genus and species of bush-cricket reared from field-collected eggs. Paraplangia sinespeculo gen. nov., sp. nov. belongs to Phaneropterinae and shares diagnostic characteristics with members of the tribe Amblycoryphini and its African subtribe Plangiina stat. nov. Paraplangia, which has a chromosome number of 31 X0, differs from other African members of the tribe and subtribe such as Eurycorypha and Plangia, which both have 29 X0. In addition to morphology, we describe the male calling song, female acoustic response, and mating behavior. As calling song, the male produces two series of short syllables. At the end of the second series the female responds with signals of similar duration and spectral composition as the male sounds (peak about 8-9 kHz). To make future identification easier, a key to all genera of Phaneropterinae found in Madagascar is presented.

}, keywords = {Amblycoryphini, bioacoustics, chromosomes, duetting, Orophus}, issn = {1082-6467}, doi = {10.3897/jor.27.24243}, url = {https://jor.pensoft.net/articles.php?id=24243}, author = {Klaus-Gerhard Heller and Claudia Hemp and Massa, Bruno and Koci{\'n}ski, Maciej and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta} } @article {52835, title = {High-speed duetting {\textendash} latency times of the female acoustic response within the bush-cricket genera Leptophyes and Andreiniimon (Orthoptera, Phaneropteridae)}, journal = {ZooKeys}, volume = {750}, year = {2018}, month = {Apr-04-2019}, pages = {45 - 58}, abstract = {

To find a mate, male and female bush-crickets of the family Phaneropteridae typically engage in duets. The male sings and the female responds. For mutual recognition, the amplitude pattern of the male song and the species-specific timing of the female response have been shown to be very important. In the seven studied species, belonging to the genera Leptophyes and Andreiniimon, these duets are extremely fast and nearly completely in the ultrasonic range. The females produce very short sounds by fast closing movements of the tegmina. They respond with species-specific delays of 20 to 150 ms after the beginning of the male song. The different latency times are probably not important for species recognition, since in sympatric species they are quite similar.

}, keywords = {duet, female acoustic signals, katydid, Phaneropterinae, stridulatory movement}, issn = {1313-2989}, doi = {10.3897/zookeys.750.23874}, url = {https://zookeys.pensoft.net/articles.php?id=23874}, author = {Klaus-Gerhard Heller and Olga S. Korsunovskaya and Massa, Bruno and Ionut Stefan Iorgu} } @article {52787, title = {Unusual abdomino-alary, defensive stridulatory mechanism in the bushcricketPantecphylus cerambycinus (Orthoptera, Tettigonioidea, Pseudophyllidae)}, journal = {Journal of Morphology}, volume = {227}, year = {1996}, month = {Jan-01-1996}, pages = {81 - 86}, issn = {0362-2525}, doi = {10.1002/(ISSN)1097-468710.1002/(SICI)1097-4687(199601)227:1<>1.0.CO;2-T10.1002/(SICI)1097-4687(199601)227:1<81::AID-JMOR6>3.0.CO;2-S}, url = {http://doi.wiley.com/10.1002/\%28ISSN\%291097-4687}, author = {Klaus-Gerhard Heller} } @article {52768, title = {Extremely divergent song types in the genus Aerotegmina Hemp (Orthoptera: Tettigoniidae: Hexacentrinae) and the description of a new species from the Eastern Arc Mountains of Tanzania (East Africa)}, journal = {Bioacoustics}, year = {2018}, month = {Feb-04-2018}, pages = {1 - 17}, abstract = {

The genus Aerotegmina comprises a group of African canopy-dwelling, predatory bushcrickets (katydids) that is known for its inflated tegmina and the unusually loud and low-frequency calling song of its type species. Here, we describe the songs of another two species of the genus which are much larger than the type species and have an even lower peak frequency. In addition, small and large species differ in song structure. While the type species and closely related forms have a broad banded, multi-peak song with strong components in the audio and ultrasonic range, the species treated here have an extremely narrow banded, resonant song with a carrier frequency of 2 kHz. With Q-values above 100 these sounds belong to the purest songs ever recorded in Orthoptera. Besides describing songs, stridulatory organs and a new species we discuss the possible functions of the song type. A key to the species of Aerotegmina is provided.

}, keywords = {Aerotegmina, bioacoustics, Eastern Arc Mountains, Orthoptera, resonant song}, issn = {0952-4622}, doi = {10.1080/09524622.2018.1443284}, url = {https://www.tandfonline.com/doi/full/10.1080/09524622.2018.1443284https://www.tandfonline.com/doi/pdf/10.1080/09524622.2018.1443284}, author = {Klaus-Gerhard Heller and Claudia Hemp} } @article {50819, title = {From an old sound recording to a new species in the genus Horatosphaga~(Orthoptera: Tettigonioidea: Phaneropterinae: Acrometopini)}, journal = {Zootaxa}, volume = {4323}, year = {2017}, month = {Oct-09-2018}, pages = {430-434}, chapter = {430}, abstract = {

Among insects, Orthoptera are a group famous for communicating by sound. This is especially true for bush-crickets (or katydids; Tettigonioidea). All acoustically active species produce sound by rubbing the fore wings (tegmina) against each other, using a stridulatory file situated on the lower side of the left tegmen and a scraper formed by the inner edge of the right tegmen (see e.g., Ragge \& Reynolds 1998). Bush-cricket species of the tribe Acrometopini often have complicated stridulatory files and also complex songs, as shown in a recent study (Hemp et al. 2017). During the preparation of that paper we discovered a sound record of a species of the group taken on 16th March 1972 by the entomologist David Hollis in Angola as part of the multi-disciplinary entomological South Western Africa Expedition (for more details see Lane et al. 2011, Pethers 2016). Hollis (principally a hemipterist) published several papers on different groups of insects (see http://bionames.org/authors/David\%20Hollis; incomplete), including Orthoptera (see Ingrisch \& Willemse 2004). The sound recording was deposited in the British Museum (Natural History) Library of Recorded Insect Sounds, and was later made openly accessible in the BioAcoustica repository (Baker et al. 2015). Hollis had found the animal at light, recorded it on the same day singing on a bush, and brought the specimen (together with two conspecific males) to the Natural History Museum (NHM), London. Here it was later identified as Horatosphaga ?stuhlmanni (Karsch 1896) by D. Ragge, the curator of Orthoptera at the NHM at that time. Under this name (Horatosphaga ?stuhlmanni) the song was described in Hemp et al. 2017. Horatosphaga males are fully winged while females are flightless and plump; sexual dimorphism reaches its extreme form in this group of species. Species of this genus are mostly recorded from open grasslands in savanna habitats up to montane elevations (e.g. H. heteromorpha, Hemp 2013).

}, issn = {1175-5326}, doi = {10.11646/zootaxa.4323.310.11646/zootaxa.4323.3.10}, url = {https://biotaxa.org/Zootaxa/issue/view/zootaxa.4323.3}, author = {Klaus-Gerhard Heller and Edward Baker} } @article {50735, title = {A new genus and other new species of Agraeciini from the Eastern Arc Mountains, East Africa (Orthoptera: Tettigoniidae; Conocephalinae; Agraeciini)}, journal = {Zootaxa}, volume = {4311}, year = {2017}, month = {Oct-08-2018}, pages = {1}, abstract = {

The genus Dendrobia n. gen. is erected to include D. amaniensis n. sp. from the East Usambara Mountains as type species. A second species in this genus, D. octopuncata n. sp., is described from the Udzungwa Mountains of Tanzania. Dendrobia n. gen. is a canopy dweller found only high up on trees in closed submontane and montane forest. The peak frequency of its song is lower than in all other known African Agraeciini, correlating with the large resonant areas in the tegmina. Two new Afroanthracites species, A. ngologolo n. sp. and A. nguru n. sp. are described from the Eastern Arc Mountains in Tanzania. These species are dwellers of understory vegetation in closed forest in the Udzungwa and Nguru Mountains respectively.

}, keywords = {bioacoustics, East Africa, Eastern Arc Mountains, new genus, new species, Orthoptera}, issn = {1175-5326}, doi = {10.11646/zootaxa.4311.110.11646/zootaxa.4311.1.1}, url = {https://biotaxa.org/Zootaxa/issue/view/zootaxa.4311.1https://biotaxa.org/Zootaxa/article/view/zootaxa.4311.1.1https://biotaxa.org/Zootaxa/article/viewFile/32748/29080https://biotaxa.org/Zootaxa/article/viewFile/32748/29080}, author = {Claudia Hemp and Ingrisch, Sigfrid and Klaus-Gerhard Heller} } @article {50695, title = {A new genus of African Acrometopini (Tettigoniidae: Phaneropterinae) based on morphology, chromosomes, acoustics, distribution, and molecular data, and the description of a new species}, journal = {Zoological Journal of the Linnean Society}, volume = {158}, year = {2010}, month = {Jan-01-2010}, pages = {66 - 82}, abstract = {

A new genus, Altihoratosphaga, is erected for species formerly assigned to Horatosphaga Schaum, 1853, and a new species is described. Four species are included in Altihoratosphaga: Altihoratosphaga nomima (Karsch, 1896), Altihoratosphaga montivaga (Sj\östedt, 1909), Altihoratosphaga nou (Hemp, 2007) and Altihoratosphaga hanangensis sp. nov. All four species are restricted to Tanzanian localities, and, except for A. nomima, for which no ecological data are available, are confined to montane forest habitats. Data on ecology, acoustics, chromosomes, and molecular relationships are provided, as well as a key to Altihoratosphaga species. The present-day distribution of Altihoratosphaga species suggests former migration events at times when wetter and colder climatic fluctuations favoured connections between montane forest communities, which today are isolated, enabling flightless taxa such as Altihoratosphaga and Monticolaria to spread.

}, keywords = {Altihoratosphaga, bush crickets, Horatosphaga, montane forest, speciation, Tanzania}, issn = {00244082}, doi = {10.1111/zoj.2010.158.issue-110.1111/j.1096-3642.2009.00542.x}, url = {http://blackwell-synergy.com/doi/abs/10.1111/zoj.2010.158.issue-1https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00542.x}, author = {Claudia Hemp and Kjetil Lysne Voje and Klaus-Gerhard Heller and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Hemp, Andreas} } @article {47981, title = {The genus Ectadia (Orthoptera: Phaneropteridae: Phaneropterinae) in East Asia: description of a new species, comparison of its complex song and duetting behavior with that of E. fulva and notes on the biology of E. fulva}, journal = {Journal of Orthoptera Research}, volume = {26}, year = {2017}, month = {Apr-06-2019}, pages = {39 - 51}, abstract = {

The genus Ectadia is an East Asian genus of the tribe Elimaeini (Phaneropterinae) and is known for the complex stridulatory files of its members. The calling song was so far known from only one species, the relatively widespread Ectadia fulva. Here a new species from Yunnan, China, is described, which is morphologically similar to E. fulva except for the stridulatory file. In this character E. fulva deviates strongly from the new species and from all other Ectadia species by its high tooth number. The long lasting song (duration 30 to 60 seconds at 25\°C) of the new species is very complex and differs widely from the much shorter song of E. fulva. Its spectral composition changes during stridulation. In the new species the females respond acoustically during gaps in the final part of the male song, whereas in E. fulva they answer typically after the male song. During mating Ectadia fulva males transfer small spermatophores without spermatophylax. All of its six nymphal stages are cryptically colored. In karyotype the new species being the first studied Ectadia species is similar to the related genera Elimaea and Ducetia (all 2n=29 chromosomes in the male).

}, keywords = {calling song, carrier frequency, male-female duet, stridulatory file, Yunnan}, issn = {1082-6467}, doi = {10.3897/jor.26.1454810.3897}, url = {https://jor.pensoft.net/articles.php?id=14548}, author = {Klaus-Gerhard Heller and Ingrisch, Sigfrid and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Chunxiang Liu} } @article {47842, title = {Warm-Up and Stridulation in the Buschcricket, Hexacentrus Unicolor Serville (Orthoptera, Conocephalidae, Listroscelidinae)}, journal = {Journal of Experimental Biology}, volume = {126}, year = {1986}, pages = {97-109}, chapter = {97}, abstract = {

The morphology of the stridulatory organ of the tropical bushcricket Hexacentrus unicolor is described and an investigation is made of stridulatory movements and muscle activities.

\ \ \  1. The song is produced by extremely fast stridulatory movements (320\–415 Hz), during which, while closing the wings, a highly specialized file is brought into contact with the plectrum to produce sound. The muscles are activated with the same frequency, i.e. they work synchronously (neurogenic).

\ \ \  2. A few minutes before the stridulation starts, the muscles are activated in a similar rhythm to that pertaining during singing (pattern of spike trains and pauses). As a result, the thorax temperature increases considerably (up to approximately 37\°C, 13\–15\°C higher than the ambient temperature). The frequency of the muscle potentials within the spike trains rises during warm-up. There are, however, indications that no phase displacement of antagonistically working muscles takes place.

}, author = {Klaus-Gerhard Heller} } @article {47758, title = {The bush-cricket Isophya kraussii (Orthoptera: Phaneropteridae): bioacoustics, distribution and description of a new subspecies from Romania}, journal = {Zootaxa}, volume = {3640}, year = {2013}, month = {May-04-2014}, pages = {258}, issn = {1175-5326}, doi = {10.11646/zootaxa.3640.210.11646/zootaxa.3640.2.8}, url = {http://biotaxa.org/Zootaxa/issue/view/zootaxa.3640.2}, author = {Ionut Stefan Iorgu and Klaus-Gerhard Heller} } @article {47513, title = {New data on the bush-cricket Montana medvedevi (Orthoptera: Tettigoniidae), critically endangered in Europe (EU 28), and a comparison of its song with all known song patterns within the genus}, journal = {Zootaxa}, volume = {4263}, year = {2017}, pages = {527-542}, chapter = {527}, abstract = {

Montana medvedevi is reported for the first time from Serbia. New information about the distribution, morphology and song of this species is discussed. The song of M. medvedevi is different from that of all other members of the genus, all figured for comparison. Montana is quite diverse regarding the amplitude pattern of the calling song of its members (known in 15 species). Surprisingly, some Montana species seem to have two song patterns, one produced during the day and one at night.

}, keywords = {bioacoustics, distribution, Platycleidini, Serbia}, doi = {10.11646/zootaxa.4263.3.5}, author = {Slobodan Ivkovi{\'c} and Ionut Stefan Iorgu and Laslo Horvat and Dragan Petrov Chobanov and Olga S. Korsunovskaya and Klaus-Gerhard Heller} } @article {47346, title = {Evolution and systematics of Green Bush-crickets (Orthoptera: Tettigoniidae: Tettigonia) in the Western Palaearctic: testing concordance between molecular, acoustic, and morphological data}, journal = {Organisms Diversity \& Evolution}, volume = {17}, year = {2017}, month = {Jan-03-2017}, pages = {213 - 228}, abstract = {

The genus Tettigonia includes 26 species distributed in the Palaearctic region. Though the Green Bush-crickets are widespread in Europe and common in a variety of habitats throughout the Palaearctic ecozone, the genus is still in need of scientific attention due to the presence of a multitude of poorly explored taxa. In the present study, we sought to clarify the evolutionary relationships of Green Bush-crickets and the composition of taxa occurring in the Western Palaearctic. Based on populations from 24 disjunct localities, the phylogeny of the group was estimated using sequences of the cytochrome oxidase subunit I (COI) and the internal transcribed spacers 1 and 2 (ITS1 and ITS2). Morphological and acoustic variation documented for the examined populations and taxa was interpreted in the context of phylogenetic relationships inferred from our genetic analyses. The trees generated in the present study supported the existence of three main lineages: \“A\”\—composed of all sampled populations of Tettigonia viridissima and the Tettigonia vaucheriana complex, \“B\”\—comprising Tettigonia caudata, Tettigonia uvarovi, and the Tettigonia armeniaca complex, and \“C\”\—consisting of Tettigonia cantans. The present study provides the first phylogenetic foundation for reviewing the systematics of Tettigonia (currently classified mostly according to morphological characteristics), proposing seven new synonymies.

}, keywords = {bioacoustics, mtDNA, phylogeny, rDNA, Tettigonia}, issn = {1439-6092}, doi = {10.1007/s13127-016-0313-3}, url = {http://link.springer.com/10.1007/s13127-016-0313-3}, author = {Grzywacz, Beata and Klaus-Gerhard Heller and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Karamysheva, Tatyana V. and Dragan Petrov Chobanov} } @article {47345, title = {Chromosomal diversification in the flightless Western Mediterranean bushcricket genus Odontura (Orthoptera: Tettigoniidae: Phaneropterinae) inferred from molecular data}, journal = {Journal of Zoological Systematics and Evolutionary Research}, volume = {52}, year = {2014}, month = {Jan-05-2014}, pages = {109 - 118}, abstract = {

We used molecular characters to infer the phylogenetic position of the Western Mediterranean bushcricket genus Odontura and to trace its high karyotype diversity. Analysis of 1391 base pairs of two mitochondrial genes (COI and ND1) and one nuclear sequence (ITS2) was conducted. Phylogenetic topologies were estimated using maximum parsimony, maximum likelihood and likelihood-based Bayesian inference. The genus Odontura is a phylogenetic outlier in respect of all other European Phaneropterinae genera and has been proposed to have originated from a hitherto unknown ancestor. Our results support the monophyly of the genus Odontura and the recognition of two subgenera: Odontura and Odonturella. We found that both Sicilian taxa of the subgenus Odontura have a completely identical morphology and song patterns. Combining these results, we proposed that both should be treated as subspecies: O. (Odontura) stenoxypha stenoxypha and O. (O.) st. arcuata. Bioacoustic data also proved to support independent markers, with song characteristics reflecting the molecular topology. Mapping the karyotypic characters onto the phylogenetic tree allows a reconstruction of the directions and transitional stages of chromosome differentiation. The number of autosomes within the genus Odontura ranges from 26 to 30. In addition to the ancestral X0 sex determination mechanism, neo-XY and neo-X1X2Y sex chromosomes have evolved independently.

}, keywords = {bioacoustics, chromosome evolution, molecular phylogeny, Odontura, Phaneropterinae}, doi = {10.1111/jzs.2014.52.issue-210.1111/jzs.12046}, url = {http://doi.wiley.com/10.1111/jzs.2014.52.issue-2}, author = {Grzywacz, Beata and Klaus-Gerhard Heller and Arne W. Lehmann and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Gerlind U.C. Lehmann} } @article {47344, title = {Review of the Plangia graminea (Serville) complex and the description of new Plangia species from East Africa (Orthoptera: Phaneropteridae, Phaneropterinae) with data on habitat, bioacoustics, and chromosomes}, journal = {Organisms Diversity \& Evolution}, volume = {15}, year = {2015}, month = {Jan-09-2015}, pages = {471 - 488}, abstract = {

The Plangia graminea complex is partly reviewed and two new species of Plangia are described, Plangia multimaculata n. sp. from savanna habitats and Plangia satiscaerulea n. sp. from the submontane zones in northern Tanzania. Plangia compressa (Walker 1869) is synonymized with P. graminea (Serville, 1838). Data on habitat, bioacoustics, and chromosomes are provided. Both analyzed Plangia species showed the same chromosomal number. Compared to other investigated African Phaneropterinae, Plangia had a reduced chromosome number and thus shows a derived condition in its genome. The sex chromosomes differed morphologically in both investigated species of Plangia suggesting different mechanisms leading to these differences. Niche specialization and the male calling song are discussed as drivers for speciation in fully alate and thus mobile taxa.

}, keywords = {bioacoustics, chromosomes, Eastern Arc Mountains, Mount Kilimanjaro, new species, speciation, Tanzania}, issn = {1439-6092}, doi = {10.1007/s13127-015-0216-8}, url = {http://link.springer.com/10.1007/s13127-015-0216-8}, author = {Claudia Hemp and Klaus-Gerhard Heller and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Grzywacz, Beata and Hemp, Andreas} } @article {47343, title = {A new genus and species of African Phaneropterinae (Orthoptera: Tettigoniidae), with data on its ecology, bioacoustics and chromosomes}, journal = {Organisms Diversity \& Evolution}, volume = {10}, year = {2010}, month = {Jan-07-2010}, pages = {215 - 226}, abstract = {

A new genus is proposed for a new East African Phaneropterinae species, Lunidia viridis, occurring on Mt. Kilimanjaro, Tanzania. Based on 33 records, notes on distribution and habitat are given, as well as acoustical data provided. Climate and vegetation parameters obtained along several transects on Mt. Kilimanjaro were evaluated describing the ecological niche of the new species. This interdisciplinary approach allows not only a profound characterisation of the ecological demands of the new genus but also predictions of the potential distribution area, which is tested for the first time for an African bush cricket species. Lunidia viridis n. gen. n. sp. occurs within humid and perhumid forests and Chagga home gardens, avoiding subhumid conditions on the mountain. It is found from 1,330 m upwards on the southern slopes, whereas the same ecological conditions are expressed from 1,930 m upwards on the drier northern slopes. Lunidia viridis has an unusually complex and variable song, which is described from field and laboratory recordings. The FISH technique for characterizing chromosomes is applied for the first time for an African species; L. viridis exhibits a karyotype typical for most Tettigoniidae.

}, keywords = {East Africa, Eastern Arc, ecological niche, Mount Kilimanjaro, new species}, issn = {1439-6092}, doi = {10.1007/s13127-010-0013-3}, url = {http://link.springer.com/10.1007/s13127-010-0013-3}, author = {Claudia Hemp and Klaus-Gerhard Heller and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Hemp, Andreas} } @article {47339, title = {The Phlesirtes complex (Orthoptera, Tettigoniidae, Conocephalinae, Conocephalini) reviewed: integrating morphological, molecular, chromosomal and bioacoustic data}, journal = {Systematic Entomology}, volume = {35}, year = {2010}, month = {Jan-07-2010}, pages = {554 - 580}, abstract = {

The tettigoniid genus Phlesirtes Bolivar and its allies are reviewed. Morphological, ecological and molecular data prompt the erection of the new genus Chortoscirtesgen.n. with type species Xiphidion meruense Sj\östedt. The genera Phlesirtes, Chortoscirtes, Karniella and Naskreckiella are characterized by morphological characters supported by molecular, acoustic, ecological and chromosomal data. Four species, Chortoscirtes pseudomeruensissp.n., C. masaicussp.n., C. puguensissp.n. and C. serengetisp.n., are described from localities in northern and coastal Tanzania and one Karniella, K. crassicercasp.n., is described from Uganda. The following comb n. are proposed: Phlesirtes kibonotensis (Sj\östedt) and Phlesirtes kilimandjaricus (Sj\östedt). Subtribal status is proposed for the four investigated African genera. A key to the Chortoscirtes species is provided.

}, doi = {10.1111/j.1365-3113.2009.00512.x}, url = {http://doi.wiley.com/10.1111/j.1365-3113.2009.00512.xhttps://api.wiley.com/onlinelibrary/tdm/v1/articles/10.1111\%2Fj.1365-3113.2009.00512.x}, author = {Claudia Hemp and Klaus-Gerhard Heller and KEHL, SIEGFRIED and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Johann Wolfgang W{\"a}gele and Hemp, Andreas} } @article {47336, title = {A new genus of African Karniellina (Orthoptera, Tettigoniidae, Conocephalinae, Conocephalini): integrating morphological, molecular and bioacoustical data}, journal = {Systematic Entomology}, volume = {35}, year = {2010}, month = {Jan-07-2010}, pages = {581 - 595}, abstract = {

Melanoscirtes gen.n. is established within Karniellina. The members of this subtribe are small conocephaline bush crickets, confined to Africa. Melanoscirtes is erected on Phlesirtes kibonotensis, a species restricted to forest clearings and forest edge in the submontane and montane zones of Mt. Kilimanjaro. A subspecies, M. kibonotensis uguenoensis, is described from the North Pare mountains, a mountain range of the Eastern Arc adjacent to Mt. Kilimanjaro. Further species of Melanoscirtes occur on other mountain ranges of the northern branch of the Eastern Arc mountains of northern Tanzania and southern Kenya. The South Pare mountains harbour M. shengenae; the West Usambaras, M. usambarensis, and the Taita Hills, M. taitensis. All species and subspecies of Melanoscirtes exhibit a similar morphology and occupy analogous habitats on the respective mountains. The song patterns for all species found within this genus are very similar, and this, together with evidence from molecular data, suggests that allopatric speciation is the reason for the biogeographic pattern found in this genus. A key for the subspecies and species of Melanoscirtes is provided.

}, doi = {10.1111/j.1365-3113.2010.00528.x}, url = {http://doi.wiley.com/10.1111/j.1365-3113.2010.00528.xhttps://api.wiley.com/onlinelibrary/tdm/v1/articles/10.1111\%2Fj.1365-3113.2010.00528.x}, author = {Claudia Hemp and KEHL, SIEGFRIED and Klaus-Gerhard Heller and Johann Wolfgang W{\"a}gele and Hemp, Andreas} } @article {47335, title = {The genus Aerotegmina (Orthoptera, Tettigoniidae, Hexacentrinae): chromosomes, morphological relations, phylogeographical patterns and description of a new species}, journal = {Organisms Diversity \& Evolution}, volume = {13}, year = {2013}, month = {Jan-12-2013}, pages = {521 - 530}, abstract = {

The genus Aerotegmina Hemp is common on East African mountains. Two species are known and a third, A. taitensis n. sp., is described in this paper. A. kilimandjarica Hemp is widespread while A. shengenae Hemp is endemic to the South Pare Mountains and A. taitensis n. sp. is known only from the Taita Hills. Morphologically, and from their song, A. shengenae and A. taitensis n. sp. are closely related. In chromosome number A. kilimandjarica (2n\ =\ 33) differs clearly from A. shengenae (2n\ =\ 27). Data presented on other flightless Orthoptera suggest that the South Pare Mountains and the Taita Hills, both belonging to the geologically old mountain chain of the Eastern Arc, show a faunistic similarity not shared by any other mountain range in the area. The mechanisms that led to this phylogeographic pattern in flightless Orthoptera in the Eastern Arc Mountains of northern Tanzania and southern Kenya and the inland volcanoes are discussed. A key to the three Aerotegmina species is presented, as well as bioacoustical data of all species compared to the phaneropterine species Euryastes jagoi.

}, keywords = {Aerotegmina, bioacoustics, chromosomes, East Africa, Eastern Arc Mountains, Euryastes, Hexacentrinae, morphology, new species, Orthoptera, phylogeography, Taita Hills}, issn = {1439-6092}, doi = {10.1007/s13127-013-0133-7}, url = {http://link.springer.com/10.1007/s13127-013-0133-7}, author = {Claudia Hemp and Klaus-Gerhard Heller and Warchalowska-Sliwa, Elzbieta and Hemp, Andreas} } @article {47334, title = {Biogeography, ecology, acoustics and chromosomes of East African Eurycorypha St{\r a}l species (Orthoptera, Phaneropterinae) with the description of new species}, journal = {Organisms Diversity \& Evolution}, volume = {13}, year = {2013}, month = {Jan-09-2013}, pages = {373 - 395}, abstract = {

Although Eurycorypha is the most species-rich Phaneropterinae genus in Africa, little is known about the distribution and the ecology of the species. In this study data on distribution, ecology, song and on chromosomes of some East African species are provided. The nymphs of some species are shown, famous for their ant-like appearance and behaviour. The male of E. punctipennis Chopard and three species of Eurycorypha are newly described. These are E. resonans n. sp. and E. combretoides n. sp. occurring in different habitats on Mount Kilimanjaro, and E. conclusa n. sp from forest habitats along the Tanzanian coast. Bioacoustically, the four recorded species are unusually divers, presenting non-resonant and resonant songs. As four Eurycorypha species occur syntopically on geological young Mount Kilimanjaro, the biogeographical pattern and possible speciation mechanisms in the genus Eurycorypha are discussed in context of the climatical history of eastern Africa.

}, keywords = {bioacoustics, Biogeography, chromosomes, ecology, Mount Kilimanjaro, new species, Orthoptera, Phaneropterinae, speciation, Tanzania}, issn = {1439-6092}, doi = {10.1007/s13127-012-0123-1}, url = {http://link.springer.com/10.1007/s13127-012-0123-1}, author = {Claudia Hemp and Klaus-Gerhard Heller and Warcha{\l}owska-{\'S}liwa, El{\.z}bieta and Grzywacz, Beata and Hemp, Andreas} } @article {40640, title = {The genus Phlesirtes Bolivar, 1922 (Orthoptera: Tettigoniidae: Conocephalinae, Conocephalini; Karniellina), a review of the genus with data on its bioacoustics and the description of new species}, journal = {Zootaxa}, volume = {4244}, year = {2017}, month = {Oct-03-2018}, pages = {451}, issn = {1175-5326}, doi = {10.11646/zootaxa.4244.410.11646/zootaxa.4244.4.1}, url = {http://biotaxa.org/Zootaxa/issue/view/zootaxa.4244.4}, author = {Claudia Hemp and Klaus-Gerhard Heller} } @book {17156, title = {Bioakustik der europ{\"a}ischen Laubheuschrecken}, series = {{\"O}kologie in Forschung und Anwendung}, volume = {1}, year = {1988}, pages = {358}, publisher = {Margraf}, organization = {Margraf}, address = {Weikersheim, Germany}, author = {Klaus-Gerhard Heller} } @article {15290, title = {Taxonomic, bioacoustic and faunistic data on a collection of Tettigonioidea from Eastern Congo (Insecta: Orthoptera)}, journal = {Zootaxa}, volume = {3785}, year = {2014}, month = {Apr-04-2014}, pages = {343}, issn = {1175-5326}, doi = {10.11646/zootaxa.3785.310.11646/zootaxa.3785.3.2}, url = {http://biotaxa.org/Zootaxa/issue/view/zootaxa.3785.3}, author = {Klaus-Gerhard Heller and Claudia Hemp and Chunxiang Liu and Volleth, Marianne} } @article {15234, title = {Surprising Genetic Diversity in Rhinolophus luctus (Chiroptera: Rhinolophidae) from Peninsular Malaysia: Description of a New Species Based on Genetic and Morphological Characters}, journal = {Acta Chiropterologica}, volume = {17}, year = {2015}, month = {Jan-06-2015}, pages = {1 - 20}, issn = {1508-1109}, doi = {10.3161/15081109ACC2015.17.1.001}, url = {http://www.bioone.org/doi/10.3161/15081109ACC2015.17.1.001}, author = {Volleth, Marianne and Loidl, Josef and Mayer, Frieder and Yong, Hoi-Sen and M{\"u}ller, Stefan and Klaus-Gerhard Heller} } @article {15202, title = {Sound production and hearing in the pyralid moth Symmoracma minoralis}, journal = {Journal of Experimental Biology}, volume = {187}, year = {1994}, pages = {101{\textendash}111}, abstract = {

Males of the moth Symmoracma minoralis (Snellen) (Lepidoptera: Pyralidae, Nymphulinae) were observed producing a high-intensity calling song (95 dB SPL at a distance of 10 cm) with a complex amplitude and frequency modulation (peaks of carrier frequency at 60 and 120 kHz). This sound is produced by a hitherto unknown type of sound organ located in the last abdominal (genital) segment, which may act as a tymbal. The observed directionality of sound output is probably achieved by means of a hollow cone surrounding the sound organ. Electrophysiological recordings revealed that the tympanal organs of S. minoralis are most sensitive in the frequency range from 50 to at least 100 kHz, which is distinctly higher than the minimum threshold levels in most other moths yet examined. The origin of genital sound production is discussed with respect to abdominal pheromone glands and pheromone-releasing movements.

}, url = {http://jeb.biologists.org/content/jexbio/187/1/101.full.pdf}, author = {Klaus-Gerhard Heller and Krahe, R{\"u}diger} } @article {15201, title = {The ultrasonic song of the moth Amyna natalis (Lepidoptera:Noctudidae: Acontiinae)}, journal = {Bioacoustics}, volume = {5}, year = {1993}, pages = {89-97}, chapter = {89}, abstract = {

In Malaysia, males of the noctuid moth Amyna natalis were observed producing a continuous ultrasonic song of high intensity (about 102 dB SPL measured at a distance of 10cm). The frequency spectrum of the sound impulses had its peak between 60 and 80 kHz. During song production the animals were perching on plants and moving their wings up and down quickly. Simultaneously, by twisting the wings it seems likely that a male-specific \“bubble\&$\#$39;\&$\#$39; in the forewing functions as a tymbal, resulting in sound production.

}, doi = {10.1080/09524622.1993.9753231}, url = {http://www.tandfonline.com/doi/abs/10.1080/09524622.1993.9753231}, author = {Klaus-Gerhard Heller and Achmann, Roland} } @article {15200, title = {Echolocation and body size in insectivorous bats: the case of the giant naked bat Cheiromeles torquatus (Molossidae)}, journal = {Le Rhinolophe}, volume = {11}, year = {1996}, pages = {27{\textendash}38}, abstract = {

Recent hypotheses propose that the maximum size of aerial insectivorous bats is limited by properties of their echolocation system. According to these hypotheses large bats would (A) have to use low call frequencies which may only allow the detection of large prey. However, large prey may be too scarce to meet a large bat\’s energy requirements. Large bats would (B) have low wing beat frequencies and due to energetic reasons also low call repetition rates. These low call repetition rates, however, many not be sufficient for prey detection. Both hypotheses were tested by analysing the echolocation behaviour of Cheiromeles torquatus (150-200g), the largest aerial insectivorous bat.

Cheriomeles searches for prey by alternating calls with the lowest frequencies at 20 and 28 kHz (call duration 30 and 24 ms respectively) at a repetition rate of 6.3Hz, and a duty cycle of 18\%. The data are discussed in relation to the foraging behaviour of Cheiromeles.

The call parameters of Cheiromeles are far removed from predictions made in both models When the data of Cheiromeles and some other large bats are included in an allometric version of model (A) the prediction of the model changes distinctly indicating that call frequency might only be a weak constraint on body size, if at all. Call repetition rate seems to be correlated to call frequency and not directly to body size as is assumed in model (B). In nay case, large bats have lower call repetition rates than wing beat rates and the difference increases with body size. Therefore other factors have to be sought to explain the size distribution of aerial insectivorous bats.

}, url = {https://www.researchgate.net/profile/Klaus-Gerhard_Heller/publication/257655672_Echolocation_and_body_size_in_insectivorous_bats_the_case_of_the_giant_naked_bat_Cheiromeles_torquatus_(Molossidae)/links/0deec52b5d412785f3000000.pdf}, author = {Klaus-Gerhard Heller} } @inbook {15183, title = {Bioacoustics}, booktitle = {Saginae of Europe}, year = {2016}, pages = {77-100}, publisher = {Edition GEEM}, organization = {Edition GEEM}, address = {Cannes}, issn = {978-2-9537533-9-4}, author = {Klaus-Gerhard Heller} } @article {15116, title = {Zur Bioakustik und Phylogenie der Gattung Poecilimon (Orthoptera, Tettigoniidae, Phaneropterinae)}, journal = {Zoologische Jahrbucher abteilung fur Systematik, okologie und geographie}, volume = {111}, year = {1984}, pages = {69-117}, chapter = {69}, abstract = {

35 European bush cricket species of the genus Poecilimon are comapred with special reference to their stridulatory movements, recorded with an opto-electronic camera, analysed in oszillograms and some power spectra, and ot their stridulatory files, studied with an electrone microscope. Since in the bush crickets the acoustics play the prime role in pair finding and thus reproductive isolation, an anslysis based on sound production and additional morphological characetristics lends itself well to investigate the relationships and the status of the species within this group. From this certain reclassifications are proposed: 15 species have been synonymized with 7 of them classified as supspecies and one subspecies has been newly described as Poecilimon obesus artedentatus.

The evolution of striduylatory movements and songs id sicussed extensively (Fig. 27). It is implied that in the songs shortneing or lengthening or fast repition of elements are typical derived forms from the original pattern. The phylogenetic tree presented in Fig. 26 considers differences in teh above physiological characteristcs as well as certian morphological onesexpecially the expression of the prothoracal stigma, whether wide or of a slit shape, the form of the pronotum, whether flat or raised, the form of the cerci, whether smooth or toothed, and finally it considers also the characteristic form of the animals stridulatory file.

}, author = {Klaus-Gerhard Heller} }