02419nas a2200229 4500008004100000022001400041245017100055210006900226260001600295300001400311490000900325520150400334653001701838653001701855653001501872653002401887653002101911100002901932700001901961700002601980856018302006 2020 eng d a1175-532600aBioacoustics of poorly known Poecilimon taxa (Insecta: Orthoptera: Tettigoniidae) with redescriptions of P. pechevi and P. stschelkanovzevi0 aBioacoustics of poorly known emPoecilimonem taxa Insecta Orthopt cApr-12-2020 a535 - 5530 v48903 a

The genus Poecilimon has been object of several studies dealing with its systematics, sound communication and evolution. Yet, published data contradict in the classification, while many taxa are still insufficiently known. In the present study we supplement the knowledge of 13 poorly known species and one additional subspecies with data on their sound communication and/or morphology. Most species concerned here are classified within two acoustic groups. First group includes P. celebi, P. obtusicercus and P. toros, showing typical characteristics of the P. syriacus group. Second group is more heterogeneous. Among that, P. pechevi, P. armeniacus, P. harveyi, P. guichardi, P. haydari, P. doga, P. davisi and P. excisus present a song pattern similar to that of P. ampliatus. In addition, P. ataturki is also quite similar in basic song structure to the latter but differs in its fine song structure. Poecilimon stschelkanovzevi, with unclear relationships, is described morphologically in detail. The present study provides hints for solving some disagreement between recent revisions, proposing unification of the Poecilimon celebi with P. syriacus group and opening discussion based on close relationships of the groups ampliatus, armeniacus, davisi, luschani and the species P. guichardi on one hand, while the complex P. ataturki/glandifer shows significant differences on the other. The acoustic peculiarities of the ampliatus group allies are discussed in an evolutionary light.

10aBarbitistini10abush-cricket10amorphology10asound communication10asound production1 aChobanov, Dragan, Petrov1 aSevgili, Hasan1 aHeller, Klaus-Gerhard uhttps://www.biotaxa.org/Zootaxa/issue/view/zootaxa.4890.4https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4890.4.6https://www.biotaxa.org/Zootaxa/article/viewFile/65904/6434600952nas a2200145 4500008004100000245014100041210007100182300001200253490000700265520042800272100002100700700001800721700002600739856004100765 2006 eng d00aPoecilimon gerlindae spec. nov.–a new bushcricket species of the Poecilimon propinquus-group (Orthoptera: Phaneropteridae) from Greece0 aPoecilimon gerlindae spec nov–a new bushcricket species of the P a109-1190 v213 a

Poecilimon gerlindae spec. nov. from Greek mainland is described. The new species is the eight known member of the P. propinquus-group (sensu Lehmann AW 1998). It is distinct with respect to male cerci, the corresponding female basal fold, body colouration, number of teeth on the stridulatory file and the num- ber of pulses in the song. Otherwise it resembles P. propinquus in body size and general song pattern.

1 aLehmann, Arne, W1 aWillemse, Fer1 aHeller, Klaus-Gerhard uhttps://bio.acousti.ca/da/node/5802802851nas a2200205 4500008004100000245012300041210006900164520208300233653001302316653001002329653002002339653001502359653003002374653003202404100001502436700002902451700002602480700002402506856011502530 2018 eng d00aReview of Poecilimon species with inflated pronotum: description of four new taxa within an acoustically diverse group0 aReview of Poecilimon species with inflated pronotum description 3 a

Poecilimon Fischer (Orthoptera, Tettigoniidae) consists of about 130 species and a functional taxonomy of the genus re- quires arranging species into phylogenetic groups. However, this is a task that requires detailed empirical testing. This paper documents data on Poecilimon species with an inflated pronotum. The Poecilimon (Poecilimon) inflatus group is a lineage ranging along a narrow zone in the south-west corner of Anatolia and in Crete plus some other Aegean islands. By producing data of morphology and calling songs of males and responding songs of females the composition and intra- group diversity of P. inflatus group are studied. Both morphology and male calling and female responding songs, exhibited a considerable intra-group diversity. In the light of data obtained the following nomenclatural changes were made. The genus Parapoecilimon Karabağ, 1975 syn. n. is synonymised with Poecilimon Fischer and Parapoecilimon antalyaensis Karabağ, 1975 is given a new combination, Poecilimon antalyaensis (Karabağ, 1975) comb. n. Four new taxa are de- scribed in the group: (i) Poecilimon isopterus sp.n.; (ii) Poecilimon inflatus lyciae subsp. n.; (iii) Poecilimon antalyaensis myrae subsp. n., and (iv) Poecilimon antalyaensis anemurium subsp. n. The Poecilimon (P.) inflatus species group con- sists of the following species: (i) Poecilimon inflatus Brunner von Wattenwyl, 1891; (ii) Poecilimon martinae Heller, 2004; (iii) Poecilimon cretensis Werner, 1903; (iv) Poecilimon antalyaensis (Karabağ, 1975) comb. n.; (v) Poecilimon bilgeri Karabağ, 1953 and (vi) Poecilimon isopterus sp.n. The species group is defined by a combination of characters including the pronotum inflated in metazona and a male cerci mostly black in their apical 1/2–1/3. The male calling song has typically two types of short syllables and the females respond acoustically with very short latency times. By using combination of morphology and song data we suggest three main lineages in the group: (i) P. bilgeri, (ii) P. antalyaensis and (iii) the other four species.

10aAnatolia10aCrete10aPhaneropterinae10aPoecilimon10aPoecilimon inflatus group10aPoecilimon isopterus sp. n.1 aKaya, Sarp1 aChobanov, Dragan, Petrov1 aHeller, Klaus-Gerhard1 aYahyaoğlu, Özgül uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4462.4https://biotaxa.org/Zootaxa/article/view/zootaxa.4462.4.102126nas a2200301 4500008004100000245017200041210006900213300000800282490000800290520119100298653001301489653001701502653001101519653002601530653001301556653002301569653002601592653002201618653001201640653001501652653002001667653001101687100002101698700002001719700002601739700001801765856004101783 2006 eng d00aMorphology, bioacoustics and phylogeography of the Isophya major group (Orthoptera: Tettigoniidae: Phaneropterinae): A species complex occurring in Anatolia and Cyprus0 aMorphology bioacoustics and phylogeography of the Isophya major  a6570 v1033 a

The species of the Isophya major-group (Orthoptera: Tettigoniidae: Phaneropterinae), are reviewed. I. major Brunner von Wattenwyl and I. mavromoustakisi Uvarov are redescribed. Two new species are described; Isophya mersinensis Sevgili & Çiplak sp. n. and Isophya salmani Sevgili & Heller sp. n. from south Anatolia. Illustrations of morphology and male calling song are provided and a detailed comparison of the four species in the group is presented. We conclude that these four species constitute a natural group in sharing male cerci with two or more denticles not ordered in a line, a character unique to this species group within the genus. From an evaluation of their morphology and song characteristics, relationships among the species in the group are I. major + (I. mersinensis sp. n. + (I. salmani + I. mavromoustakisi). Based on the the distribution pattern and habitat preference, we concluded that the division of I. salmani and I. mavromoustakisi was a vicariant event resulting from the separation of Cyprus and Anatolia due to reflooding of the Mediterranean after the Messinian salinity crisis at the beginning of the Pliocene, around 5 My ago.

10aAnatolia10aBiogeography10aCyprus10aEastern Mediterranean10aI. major10aI. mavromoustakisi10aI. mersinensis sp. n.10aI. salmani sp. n.10aIsophya10aOrthoptera10aPhaneropterinae10aTurkey1 aSevgīlī, Hasan1 aÇiplak, Battal1 aHeller, Klaus-Gerhard1 aDemirsoy, Ali uhttps://bio.acousti.ca/da/node/5794200606nas a2200169 4500008004100000022001400041245012300055210006900178260001600247300000800263490000900271100001500280700002900295700002600324700002400350856006200374 2018 eng d a1175-532600aReview of Poecilimon species with inflated pronotum: description of four new taxa within an acoustically diverse group0 aReview of Poecilimon species with inflated pronotum description  cMay-08-2020 a4510 v44621 aKaya, Sarp1 aChobanov, Dragan, Petrov1 aHeller, Klaus-Gerhard1 aYahyaoğlu, Özgül uhttps://biotaxa.org/Zootaxa/article/view/zootaxa.4462.4.101905nas a2200253 4500008004100000022001400041245019100055210006900246260001600315300001400331490000700345520105300352653001901405653001701424653001601441653001301457653001201470100002601482700001801508700001701526700002201543700003601565856005001601 2018 eng d a1082-646700aParaplangia sinespeculo, a new genus and species of bush-cricket, with notes on its biology and a key to the genera of Phaneropterinae (Orthoptera: Tettigonioidea) from Madagascar0 aiParaplangia sinespeculoi a new genus and species of bushcricket cNov-07-2019 a143 - 1530 v273 a

Madagascar is a well-known hotspot of biodiversity. However, many Orthoptera, and especially the Tettigonioidea, belong to little-studied groups. Here we describe a new genus and species of bush-cricket reared from field-collected eggs. Paraplangia sinespeculo gen. nov., sp. nov. belongs to Phaneropterinae and shares diagnostic characteristics with members of the tribe Amblycoryphini and its African subtribe Plangiina stat. nov. Paraplangia, which has a chromosome number of 31 X0, differs from other African members of the tribe and subtribe such as Eurycorypha and Plangia, which both have 29 X0. In addition to morphology, we describe the male calling song, female acoustic response, and mating behavior. As calling song, the male produces two series of short syllables. At the end of the second series the female responds with signals of similar duration and spectral composition as the male sounds (peak about 8-9 kHz). To make future identification easier, a key to all genera of Phaneropterinae found in Madagascar is presented.

10aAmblycoryphini10abioacoustics10achromosomes10aduetting10aOrophus1 aHeller, Klaus-Gerhard1 aHemp, Claudia1 aMassa, Bruno1 aKociński, Maciej1 aWarchałowska-Śliwa, Elżbieta uhttps://jor.pensoft.net/articles.php?id=2424301592nas a2200241 4500008004100000022001400041245016300055210006900218260001600287300001200303490000800315520078000323653000901103653002801112653001201140653002001152653002601172100002601198700002801224700001701252700002701269856005401296 2018 eng d a1313-298900aHigh-speed duetting – latency times of the female acoustic response within the bush-cricket genera Leptophyes and Andreiniimon (Orthoptera, Phaneropteridae)0 aHighspeed duetting latency times of the female acoustic response cApr-04-2019 a45 - 580 v7503 a

To find a mate, male and female bush-crickets of the family Phaneropteridae typically engage in duets. The male sings and the female responds. For mutual recognition, the amplitude pattern of the male song and the species-specific timing of the female response have been shown to be very important. In the seven studied species, belonging to the genera Leptophyes and Andreiniimon, these duets are extremely fast and nearly completely in the ultrasonic range. The females produce very short sounds by fast closing movements of the tegmina. They respond with species-specific delays of 20 to 150 ms after the beginning of the male song. The different latency times are probably not important for species recognition, since in sympatric species they are quite similar.

10aduet10afemale acoustic signals10akatydid10aPhaneropterinae10astridulatory movement1 aHeller, Klaus-Gerhard1 aKorsunovskaya, Olga, S.1 aMassa, Bruno1 aIorgu, Ionuț, Ștefan uhttps://zookeys.pensoft.net/articles.php?id=2387400524nas a2200133 4500008004100000022001400041245015100055210006900206260001600275300001200291490000800303100002600311856005300337 1996 eng d a0362-252500aUnusual abdomino-alary, defensive stridulatory mechanism in the bushcricketPantecphylus cerambycinus (Orthoptera, Tettigonioidea, Pseudophyllidae)0 aUnusual abdominoalary defensive stridulatory mechanism in the bu cJan-01-1996 a81 - 860 v2271 aHeller, Klaus-Gerhard uhttp://doi.wiley.com/10.1002/%28ISSN%291097-468701786nas a2200205 4500008004100000022001400041245020900055210006900264260001600333300001100349520094800360653001601308653001701324653002601341653001501367653001801382100002601400700001801426856013601444 2018 eng d a0952-462200aExtremely divergent song types in the genus Aerotegmina Hemp (Orthoptera: Tettigoniidae: Hexacentrinae) and the description of a new species from the Eastern Arc Mountains of Tanzania (East Africa)0 aExtremely divergent song types in the genus iAerotegminai Hemp O cFeb-04-2018 a1 - 173 a

The genus Aerotegmina comprises a group of African canopy-dwelling, predatory bushcrickets (katydids) that is known for its inflated tegmina and the unusually loud and low-frequency calling song of its type species. Here, we describe the songs of another two species of the genus which are much larger than the type species and have an even lower peak frequency. In addition, small and large species differ in song structure. While the type species and closely related forms have a broad banded, multi-peak song with strong components in the audio and ultrasonic range, the species treated here have an extremely narrow banded, resonant song with a carrier frequency of 2 kHz. With Q-values above 100 these sounds belong to the purest songs ever recorded in Orthoptera. Besides describing songs, stridulatory organs and a new species we discuss the possible functions of the song type. A key to the species of Aerotegmina is provided.

10aAerotegmina10abioacoustics10aEastern Arc Mountains10aOrthoptera10aresonant song1 aHeller, Klaus-Gerhard1 aHemp, Claudia uhttps://www.tandfonline.com/doi/full/10.1080/09524622.2018.1443284https://www.tandfonline.com/doi/pdf/10.1080/09524622.2018.144328402561nas a2200157 4500008004100000022001400041245014300055210006900198260001600267300001200283490000900295520199700304100002602301700001802327856005802345 2017 eng d a1175-532600aFrom an old sound recording to a new species in the genus Horatosphaga (Orthoptera: Tettigonioidea: Phaneropterinae: Acrometopini)0 aFrom an old sound recording to a new species in the genus iHorat cOct-09-2018 a430-4340 v43233 a

Among insects, Orthoptera are a group famous for communicating by sound. This is especially true for bush-crickets (or katydids; Tettigonioidea). All acoustically active species produce sound by rubbing the fore wings (tegmina) against each other, using a stridulatory file situated on the lower side of the left tegmen and a scraper formed by the inner edge of the right tegmen (see e.g., Ragge & Reynolds 1998). Bush-cricket species of the tribe Acrometopini often have complicated stridulatory files and also complex songs, as shown in a recent study (Hemp et al. 2017). During the preparation of that paper we discovered a sound record of a species of the group taken on 16th March 1972 by the entomologist David Hollis in Angola as part of the multi-disciplinary entomological South Western Africa Expedition (for more details see Lane et al. 2011, Pethers 2016). Hollis (principally a hemipterist) published several papers on different groups of insects (see http://bionames.org/authors/David%20Hollis; incomplete), including Orthoptera (see Ingrisch & Willemse 2004). The sound recording was deposited in the British Museum (Natural History) Library of Recorded Insect Sounds, and was later made openly accessible in the BioAcoustica repository (Baker et al. 2015). Hollis had found the animal at light, recorded it on the same day singing on a bush, and brought the specimen (together with two conspecific males) to the Natural History Museum (NHM), London. Here it was later identified as Horatosphaga ?stuhlmanni (Karsch 1896) by D. Ragge, the curator of Orthoptera at the NHM at that time. Under this name (Horatosphaga ?stuhlmanni) the song was described in Hemp et al. 2017. Horatosphaga males are fully winged while females are flightless and plump; sexual dimorphism reaches its extreme form in this group of species. Species of this genus are mostly recorded from open grasslands in savanna habitats up to montane elevations (e.g. H. heteromorpha, Hemp 2013).

1 aHeller, Klaus-Gerhard1 aBaker, Edward uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4323.301688nas a2200241 4500008004100000022001400041245015200055210006900207260001600276300000600292490000900298520074200307653001701049653001601066653002601082653001401108653001601122653001501138100001801153700002201171700002601193856022701219 2017 eng d a1175-532600aA new genus and other new species of Agraeciini from the Eastern Arc Mountains, East Africa (Orthoptera: Tettigoniidae; Conocephalinae; Agraeciini)0 anew genus and other new species of Agraeciini from the Eastern A cOct-08-2018 a10 v43113 a

The genus Dendrobia n. gen. is erected to include D. amaniensis n. sp. from the East Usambara Mountains as type species. A second species in this genus, D. octopuncata n. sp., is described from the Udzungwa Mountains of Tanzania. Dendrobia n. gen. is a canopy dweller found only high up on trees in closed submontane and montane forest. The peak frequency of its song is lower than in all other known African Agraeciini, correlating with the large resonant areas in the tegmina. Two new Afroanthracites species, A. ngologolo n. sp. and A. nguru n. sp. are described from the Eastern Arc Mountains in Tanzania. These species are dwellers of understory vegetation in closed forest in the Udzungwa and Nguru Mountains respectively.

10abioacoustics10aEast Africa10aEastern Arc Mountains10anew genus10anew species10aOrthoptera1 aHemp, Claudia1 aIngrisch, Sigfrid1 aHeller, Klaus-Gerhard uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4311.1https://biotaxa.org/Zootaxa/article/view/zootaxa.4311.1.1https://biotaxa.org/Zootaxa/article/viewFile/32748/29080https://biotaxa.org/Zootaxa/article/viewFile/32748/2908001959nas a2200265 4500008004100000022001300041245018900054210006900243260001600312300001200328490000800340520096300348653002101311653001801332653001701350653001901367653001501386653001301401100001801414700002401432700002601456700003601482700001801518856015701536 2010 eng d a0024408200aA new genus of African Acrometopini (Tettigoniidae: Phaneropterinae) based on morphology, chromosomes, acoustics, distribution, and molecular data, and the description of a new species0 anew genus of African Acrometopini Tettigoniidae Phaneropterinae  cJan-01-2010 a66 - 820 v1583 a

A new genus, Altihoratosphaga, is erected for species formerly assigned to Horatosphaga Schaum, 1853, and a new species is described. Four species are included in Altihoratosphaga: Altihoratosphaga nomima (Karsch, 1896), Altihoratosphaga montivaga (Sjöstedt, 1909), Altihoratosphaga nou (Hemp, 2007) and Altihoratosphaga hanangensis sp. nov. All four species are restricted to Tanzanian localities, and, except for A. nomima, for which no ecological data are available, are confined to montane forest habitats. Data on ecology, acoustics, chromosomes, and molecular relationships are provided, as well as a key to Altihoratosphaga species. The present-day distribution of Altihoratosphaga species suggests former migration events at times when wetter and colder climatic fluctuations favoured connections between montane forest communities, which today are isolated, enabling flightless taxa such as Altihoratosphaga and Monticolaria to spread.

10aAltihoratosphaga10abush crickets10aHoratosphaga10amontane forest10aspeciation10aTanzania1 aHemp, Claudia1 aVoje, Kjetil, Lysne1 aHeller, Klaus-Gerhard1 aWarchałowska-Śliwa, Elżbieta1 aHemp, Andreas uhttp://blackwell-synergy.com/doi/abs/10.1111/zoj.2010.158.issue-1https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00542.x02084nas a2200241 4500008004100000022001400041245024600055210006900301260001600370300001200386490000700398520119100405653001701596653002201613653002101635653002201656653001101678100002601689700002201715700003601737700001901773856005001792 2017 eng d a1082-646700aThe genus Ectadia (Orthoptera: Phaneropteridae: Phaneropterinae) in East Asia: description of a new species, comparison of its complex song and duetting behavior with that of E. fulva and notes on the biology of E. fulva0 agenus iEctadiai Orthoptera Phaneropteridae Phaneropterinae in Ea cApr-06-2019 a39 - 510 v263 a

The genus Ectadia is an East Asian genus of the tribe Elimaeini (Phaneropterinae) and is known for the complex stridulatory files of its members. The calling song was so far known from only one species, the relatively widespread Ectadia fulva. Here a new species from Yunnan, China, is described, which is morphologically similar to E. fulva except for the stridulatory file. In this character E. fulva deviates strongly from the new species and from all other Ectadia species by its high tooth number. The long lasting song (duration 30 to 60 seconds at 25°C) of the new species is very complex and differs widely from the much shorter song of E. fulva. Its spectral composition changes during stridulation. In the new species the females respond acoustically during gaps in the final part of the male song, whereas in E. fulva they answer typically after the male song. During mating Ectadia fulva males transfer small spermatophores without spermatophylax. All of its six nymphal stages are cryptically colored. In karyotype the new species being the first studied Ectadia species is similar to the related genera Elimaea and Ducetia (all 2n=29 chromosomes in the male).

10acalling song10acarrier frequency10amale-female duet10astridulatory file10aYunnan1 aHeller, Klaus-Gerhard1 aIngrisch, Sigfrid1 aWarchałowska-Śliwa, Elżbieta1 aLiu, Chunxiang uhttps://jor.pensoft.net/articles.php?id=1454801503nas a2200121 4500008004100000245012700041210006900168300001100237490000800248520105800256100002601314856004101340 1986 eng d00aWarm-Up and Stridulation in the Buschcricket, Hexacentrus Unicolor Serville (Orthoptera, Conocephalidae, Listroscelidinae)0 aWarmUp and Stridulation in the Buschcricket Hexacentrus Unicolor a97-1090 v1263 a

The morphology of the stridulatory organ of the tropical bushcricket Hexacentrus unicolor is described and an investigation is made of stridulatory movements and muscle activities.

    1. The song is produced by extremely fast stridulatory movements (320–415 Hz), during which, while closing the wings, a highly specialized file is brought into contact with the plectrum to produce sound. The muscles are activated with the same frequency, i.e. they work synchronously (neurogenic).

    2. A few minutes before the stridulation starts, the muscles are activated in a similar rhythm to that pertaining during singing (pattern of spike trains and pauses). As a result, the thorax temperature increases considerably (up to approximately 37°C, 13–15°C higher than the ambient temperature). The frequency of the muscle potentials within the spike trains rises during warm-up. There are, however, indications that no phase displacement of antagonistically working muscles takes place.

1 aHeller, Klaus-Gerhard uhttps://bio.acousti.ca/da/node/4784200565nas a2200145 4500008004100000022001400041245015200055210006900207260001600276300000800292490000900300100002700309700002600336856005700362 2013 eng d a1175-532600aThe bush-cricket Isophya kraussii (Orthoptera: Phaneropteridae): bioacoustics, distribution and description of a new subspecies from Romania0 abushcricket iIsophya kraussiii Orthoptera Phaneropteridae bioaco cMay-04-2014 a2580 v36401 aIorgu, Ionuț, Ștefan1 aHeller, Klaus-Gerhard uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.3640.201316nas a2200229 4500008004100000245019600041210006900237300001200306490000900318520050400327653001700831653001700848653001800865653001100883100002300894700002700917700001800944700002900962700002800991700002601019856004101045 2017 eng d00aNew data on the bush-cricket Montana medvedevi (Orthoptera: Tettigoniidae), critically endangered in Europe (EU 28), and a comparison of its song with all known song patterns within the genus0 aNew data on the bushcricket Montana medvedevi Orthoptera Tettigo a527-5420 v42633 a

Montana medvedevi is reported for the first time from Serbia. New information about the distribution, morphology and song of this species is discussed. The song of M. medvedevi is different from that of all other members of the genus, all figured for comparison. Montana is quite diverse regarding the amplitude pattern of the calling song of its members (known in 15 species). Surprisingly, some Montana species seem to have two song patterns, one produced during the day and one at night.

10abioacoustics10adistribution10aPlatycleidini10aSerbia1 aIvković, Slobodan1 aIorgu, Ionuț, Ștefan1 aHorvat, Laslo1 aChobanov, Dragan, Petrov1 aKorsunovskaya, Olga, S.1 aHeller, Klaus-Gerhard uhttps://bio.acousti.ca/da/node/4751302352nas a2200253 4500008004100000022001400041245019300055210006900248260001600317300001400333490000700347520148400354653001701838653001001855653001401865653000901879653001501888100002001903700002601923700003601949700002901985700002902014856005502043 2017 eng d a1439-609200aEvolution and systematics of Green Bush-crickets (Orthoptera: Tettigoniidae: Tettigonia) in the Western Palaearctic: testing concordance between molecular, acoustic, and morphological data0 aEvolution and systematics of Green Bushcrickets Orthoptera Tetti cJan-03-2017 a213 - 2280 v173 a

The genus Tettigonia includes 26 species distributed in the Palaearctic region. Though the Green Bush-crickets are widespread in Europe and common in a variety of habitats throughout the Palaearctic ecozone, the genus is still in need of scientific attention due to the presence of a multitude of poorly explored taxa. In the present study, we sought to clarify the evolutionary relationships of Green Bush-crickets and the composition of taxa occurring in the Western Palaearctic. Based on populations from 24 disjunct localities, the phylogeny of the group was estimated using sequences of the cytochrome oxidase subunit I (COI) and the internal transcribed spacers 1 and 2 (ITS1 and ITS2). Morphological and acoustic variation documented for the examined populations and taxa was interpreted in the context of phylogenetic relationships inferred from our genetic analyses. The trees generated in the present study supported the existence of three main lineages: “A”—composed of all sampled populations of Tettigonia viridissima and the Tettigonia vaucheriana complex, “B”—comprising Tettigonia caudata, Tettigonia uvarovi, and the Tettigonia armeniaca complex, and “C”—consisting of Tettigonia cantans. The present study provides the first phylogenetic foundation for reviewing the systematics of Tettigonia (currently classified mostly according to morphological characteristics), proposing seven new synonymies.

10abioacoustics10amtDNA10aphylogeny10arDNA10aTettigonia1 aGrzywacz, Beata1 aHeller, Klaus-Gerhard1 aWarchałowska-Śliwa, Elżbieta1 aKaramysheva, Tatyana, V.1 aChobanov, Dragan, Petrov uhttp://link.springer.com/10.1007/s13127-016-0313-302299nas a2200241 4500008004100000245018000041210006900221260001600290300001400306490000700320520144700327653001701774653002501791653002401816653001301840653002001853100002001873700002601893700002201919700003601941700002701977856005302004 2014 eng d00aChromosomal diversification in the flightless Western Mediterranean bushcricket genus Odontura (Orthoptera: Tettigoniidae: Phaneropterinae) inferred from molecular data0 aChromosomal diversification in the flightless Western Mediterran cJan-05-2014 a109 - 1180 v523 a

We used molecular characters to infer the phylogenetic position of the Western Mediterranean bushcricket genus Odontura and to trace its high karyotype diversity. Analysis of 1391 base pairs of two mitochondrial genes (COI and ND1) and one nuclear sequence (ITS2) was conducted. Phylogenetic topologies were estimated using maximum parsimony, maximum likelihood and likelihood-based Bayesian inference. The genus Odontura is a phylogenetic outlier in respect of all other European Phaneropterinae genera and has been proposed to have originated from a hitherto unknown ancestor. Our results support the monophyly of the genus Odontura and the recognition of two subgenera: Odontura and Odonturella. We found that both Sicilian taxa of the subgenus Odontura have a completely identical morphology and song patterns. Combining these results, we proposed that both should be treated as subspecies: O. (Odontura) stenoxypha stenoxypha and O. (O.) st. arcuata. Bioacoustic data also proved to support independent markers, with song characteristics reflecting the molecular topology. Mapping the karyotypic characters onto the phylogenetic tree allows a reconstruction of the directions and transitional stages of chromosome differentiation. The number of autosomes within the genus Odontura ranges from 26 to 30. In addition to the ancestral X0 sex determination mechanism, neo-XY and neo-X1X2Y sex chromosomes have evolved independently.

10abioacoustics10achromosome evolution10amolecular phylogeny10aOdontura10aPhaneropterinae1 aGrzywacz, Beata1 aHeller, Klaus-Gerhard1 aLehmann, Arne, W.1 aWarchałowska-Śliwa, Elżbieta1 aLehmann, Gerlind, U.C. uhttp://doi.wiley.com/10.1111/jzs.2014.52.issue-201815nas a2200277 4500008004100000022001400041245021300055210006900268260001600337300001400353490000700367520086500374653001701239653001601256653002601272653002201298653001601320653001501336653001301351100001801364700002601382700003601408700002001444700001801464856005501482 2015 eng d a1439-609200aReview of the Plangia graminea (Serville) complex and the description of new Plangia species from East Africa (Orthoptera: Phaneropteridae, Phaneropterinae) with data on habitat, bioacoustics, and chromosomes0 aReview of the Plangia graminea Serville complex and the descript cJan-09-2015 a471 - 4880 v153 a

The Plangia graminea complex is partly reviewed and two new species of Plangia are described, Plangia multimaculata n. sp. from savanna habitats and Plangia satiscaerulea n. sp. from the submontane zones in northern Tanzania. Plangia compressa (Walker 1869) is synonymized with P. graminea (Serville, 1838). Data on habitat, bioacoustics, and chromosomes are provided. Both analyzed Plangia species showed the same chromosomal number. Compared to other investigated African Phaneropterinae, Plangia had a reduced chromosome number and thus shows a derived condition in its genome. The sex chromosomes differed morphologically in both investigated species of Plangia suggesting different mechanisms leading to these differences. Niche specialization and the male calling song are discussed as drivers for speciation in fully alate and thus mobile taxa.

10abioacoustics10achromosomes10aEastern Arc Mountains10aMount Kilimanjaro10anew species10aspeciation10aTanzania1 aHemp, Claudia1 aHeller, Klaus-Gerhard1 aWarchałowska-Śliwa, Elżbieta1 aGrzywacz, Beata1 aHemp, Andreas uhttp://link.springer.com/10.1007/s13127-015-0216-802021nas a2200241 4500008004100000022001400041245013900055210006900194260001600263300001400279490000700293520123500300653001601535653001601551653002101567653002201588653001601610100001801626700002601644700003601670700001801706856005501724 2010 eng d a1439-609200aA new genus and species of African Phaneropterinae (Orthoptera: Tettigoniidae), with data on its ecology, bioacoustics and chromosomes0 anew genus and species of African Phaneropterinae Orthoptera Tett cJan-07-2010 a215 - 2260 v103 a

A new genus is proposed for a new East African Phaneropterinae species, Lunidia viridis, occurring on Mt. Kilimanjaro, Tanzania. Based on 33 records, notes on distribution and habitat are given, as well as acoustical data provided. Climate and vegetation parameters obtained along several transects on Mt. Kilimanjaro were evaluated describing the ecological niche of the new species. This interdisciplinary approach allows not only a profound characterisation of the ecological demands of the new genus but also predictions of the potential distribution area, which is tested for the first time for an African bush cricket species. Lunidia viridis n. gen. n. sp. occurs within humid and perhumid forests and Chagga home gardens, avoiding subhumid conditions on the mountain. It is found from 1,330 m upwards on the southern slopes, whereas the same ecological conditions are expressed from 1,930 m upwards on the drier northern slopes. Lunidia viridis has an unusually complex and variable song, which is described from field and laboratory recordings. The FISH technique for characterizing chromosomes is applied for the first time for an African species; L. viridis exhibits a karyotype typical for most Tettigoniidae.

10aEast Africa10aEastern Arc10aecological niche10aMount Kilimanjaro10anew species1 aHemp, Claudia1 aHeller, Klaus-Gerhard1 aWarchałowska-Śliwa, Elżbieta1 aHemp, Andreas uhttp://link.springer.com/10.1007/s13127-010-0013-301697nas a2200193 4500008004100000245016700041210006900208260001600277300001400293490000700307520089700314100001801211700002601229700002001255700003601275700003001311700001801341856014401359 2010 eng d00aThe Phlesirtes complex (Orthoptera, Tettigoniidae, Conocephalinae, Conocephalini) reviewed: integrating morphological, molecular, chromosomal and bioacoustic data0 aPhlesirtes complex Orthoptera Tettigoniidae Conocephalinae Conoc cJan-07-2010 a554 - 5800 v353 a

The tettigoniid genus Phlesirtes Bolivar and its allies are reviewed. Morphological, ecological and molecular data prompt the erection of the new genus Chortoscirtesgen.n. with type species Xiphidion meruense Sjöstedt. The genera Phlesirtes, Chortoscirtes, Karniella and Naskreckiella are characterized by morphological characters supported by molecular, acoustic, ecological and chromosomal data. Four species, Chortoscirtes pseudomeruensissp.n., C. masaicussp.n., C. puguensissp.n. and C. serengetisp.n., are described from localities in northern and coastal Tanzania and one Karniella, K. crassicercasp.n., is described from Uganda. The following comb n. are proposed: Phlesirtes kibonotensis (Sjöstedt) and Phlesirtes kilimandjaricus (Sjöstedt). Subtribal status is proposed for the four investigated African genera. A key to the Chortoscirtes species is provided.

1 aHemp, Claudia1 aHeller, Klaus-Gerhard1 aKEHL, SIEGFRIED1 aWarchałowska-Śliwa, Elżbieta1 aWägele, Johann, Wolfgang1 aHemp, Andreas uhttp://doi.wiley.com/10.1111/j.1365-3113.2009.00512.xhttps://api.wiley.com/onlinelibrary/tdm/v1/articles/10.1111%2Fj.1365-3113.2009.00512.x01925nas a2200181 4500008004100000245015900041210006900200260001600269300001400285490000700299520118100306100001801487700002001505700002601525700003001551700001801581856014401599 2010 eng d00aA new genus of African Karniellina (Orthoptera, Tettigoniidae, Conocephalinae, Conocephalini): integrating morphological, molecular and bioacoustical data0 anew genus of African Karniellina Orthoptera Tettigoniidae Conoce cJan-07-2010 a581 - 5950 v353 a

Melanoscirtes gen.n. is established within Karniellina. The members of this subtribe are small conocephaline bush crickets, confined to Africa. Melanoscirtes is erected on Phlesirtes kibonotensis, a species restricted to forest clearings and forest edge in the submontane and montane zones of Mt. Kilimanjaro. A subspecies, M. kibonotensis uguenoensis, is described from the North Pare mountains, a mountain range of the Eastern Arc adjacent to Mt. Kilimanjaro. Further species of Melanoscirtes occur on other mountain ranges of the northern branch of the Eastern Arc mountains of northern Tanzania and southern Kenya. The South Pare mountains harbour M. shengenae; the West Usambaras, M. usambarensis, and the Taita Hills, M. taitensis. All species and subspecies of Melanoscirtes exhibit a similar morphology and occupy analogous habitats on the respective mountains. The song patterns for all species found within this genus are very similar, and this, together with evidence from molecular data, suggests that allopatric speciation is the reason for the biogeographic pattern found in this genus. A key for the subspecies and species of Melanoscirtes is provided.

1 aHemp, Claudia1 aKEHL, SIEGFRIED1 aHeller, Klaus-Gerhard1 aWägele, Johann, Wolfgang1 aHemp, Andreas uhttp://doi.wiley.com/10.1111/j.1365-3113.2010.00528.xhttps://api.wiley.com/onlinelibrary/tdm/v1/articles/10.1111%2Fj.1365-3113.2010.00528.x02151nas a2200325 4500008004100000022001400041245016800055210006900223260001600292300001400308490000700322520114200329653001601471653001701487653001601504653001601520653002601536653001401562653001801576653001501594653001601609653001501625653001901640653001601659100001801675700002601693700003301719700001801752856005501770 2013 eng d a1439-609200aThe genus Aerotegmina (Orthoptera, Tettigoniidae, Hexacentrinae): chromosomes, morphological relations, phylogeographical patterns and description of a new species0 agenus Aerotegmina Orthoptera Tettigoniidae Hexacentrinae chromos cJan-12-2013 a521 - 5300 v133 a

The genus Aerotegmina Hemp is common on East African mountains. Two species are known and a third, A. taitensis n. sp., is described in this paper. A. kilimandjarica Hemp is widespread while A. shengenae Hemp is endemic to the South Pare Mountains and A. taitensis n. sp. is known only from the Taita Hills. Morphologically, and from their song, A. shengenae and A. taitensis n. sp. are closely related. In chromosome number A. kilimandjarica (2n = 33) differs clearly from A. shengenae (2n = 27). Data presented on other flightless Orthoptera suggest that the South Pare Mountains and the Taita Hills, both belonging to the geologically old mountain chain of the Eastern Arc, show a faunistic similarity not shared by any other mountain range in the area. The mechanisms that led to this phylogeographic pattern in flightless Orthoptera in the Eastern Arc Mountains of northern Tanzania and southern Kenya and the inland volcanoes are discussed. A key to the three Aerotegmina species is presented, as well as bioacoustical data of all species compared to the phaneropterine species Euryastes jagoi.

10aAerotegmina10abioacoustics10achromosomes10aEast Africa10aEastern Arc Mountains10aEuryastes10aHexacentrinae10amorphology10anew species10aOrthoptera10aphylogeography10aTaita Hills1 aHemp, Claudia1 aHeller, Klaus-Gerhard1 aWarchalowska-Sliwa, Elzbieta1 aHemp, Andreas uhttp://link.springer.com/10.1007/s13127-013-0133-701960nas a2200313 4500008004100000022001400041245016100055210006900216260001600285300001400301490000700315520098800322653001701310653001701327653001601344653001201360653002201372653001601394653001501410653002001425653001501445653001301460100001801473700002601491700003601517700002001553700001801573856005501591 2013 eng d a1439-609200aBiogeography, ecology, acoustics and chromosomes of East African Eurycorypha Stål species (Orthoptera, Phaneropterinae) with the description of new species0 aBiogeography ecology acoustics and chromosomes of East African E cJan-09-2013 a373 - 3950 v133 a

Although Eurycorypha is the most species-rich Phaneropterinae genus in Africa, little is known about the distribution and the ecology of the species. In this study data on distribution, ecology, song and on chromosomes of some East African species are provided. The nymphs of some species are shown, famous for their ant-like appearance and behaviour. The male of E. punctipennis Chopard and three species of Eurycorypha are newly described. These are E. resonans n. sp. and E. combretoides n. sp. occurring in different habitats on Mount Kilimanjaro, and E. conclusa n. sp from forest habitats along the Tanzanian coast. Bioacoustically, the four recorded species are unusually divers, presenting non-resonant and resonant songs. As four Eurycorypha species occur syntopically on geological young Mount Kilimanjaro, the biogeographical pattern and possible speciation mechanisms in the genus Eurycorypha are discussed in context of the climatical history of eastern Africa.

10abioacoustics10aBiogeography10achromosomes10aecology10aMount Kilimanjaro10anew species10aOrthoptera10aPhaneropterinae10aspeciation10aTanzania1 aHemp, Claudia1 aHeller, Klaus-Gerhard1 aWarchałowska-Śliwa, Elżbieta1 aGrzywacz, Beata1 aHemp, Andreas uhttp://link.springer.com/10.1007/s13127-012-0123-100603nas a2200145 4500008004100000022001400041245019900055210006900254260001600323300000800339490000900347100001800356700002600374856005700400 2017 eng d a1175-532600aThe genus Phlesirtes Bolivar, 1922 (Orthoptera: Tettigoniidae: Conocephalinae, Conocephalini; Karniellina), a review of the genus with data on its bioacoustics and the description of new species0 agenus Phlesirtes Bolivar 1922 Orthoptera Tettigoniidae Conocepha cOct-03-2018 a4510 v42441 aHemp, Claudia1 aHeller, Klaus-Gerhard uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.4244.400378nam a2200121 4500008004100000245005000041210005000091260003400141300000800175490000600183100002600189856004100215 1988 eng d00aBioakustik der europäischen Laubheuschrecken0 aBioakustik der europäischen Laubheuschrecken aWeikersheim, GermanybMargraf a3580 v11 aHeller, Klaus-Gerhard uhttps://bio.acousti.ca/da/node/1715600590nas a2200169 4500008004100000022001400041245012100055210006900176260001600245300000800261490000900269100002600278700001800304700001900322700002200341856005700363 2014 eng d a1175-532600aTaxonomic, bioacoustic and faunistic data on a collection of Tettigonioidea from Eastern Congo (Insecta: Orthoptera)0 aTaxonomic bioacoustic and faunistic data on a collection of Tett cApr-04-2014 a3430 v37851 aHeller, Klaus-Gerhard1 aHemp, Claudia1 aLiu, Chunxiang1 aVolleth, Marianne uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.3785.300721nas a2200193 4500008004100000022001400041245018400055210006900239260001600308300001100324490000700335100002200342700001700364700001900381700001800400700002000418700002600438856006300464 2015 eng d a1508-110900aSurprising Genetic Diversity in Rhinolophus luctus (Chiroptera: Rhinolophidae) from Peninsular Malaysia: Description of a New Species Based on Genetic and Morphological Characters0 aSurprising Genetic Diversity in Rhinolophus luctus Chiroptera Rh cJan-06-2015 a1 - 200 v171 aVolleth, Marianne1 aLoidl, Josef1 aMayer, Frieder1 aYong, Hoi-Sen1 aMüller, Stefan1 aHeller, Klaus-Gerhard uhttp://www.bioone.org/doi/10.3161/15081109ACC2015.17.1.00101376nas a2200133 4500008004100000245007400041210006900115300001400184490000800198520092600206100002601132700002001158856006401178 1994 eng d00aSound production and hearing in the pyralid moth Symmoracma minoralis0 aSound production and hearing in the pyralid moth Symmoracma mino a101–1110 v1873 a

Males of the moth Symmoracma minoralis (Snellen) (Lepidoptera: Pyralidae, Nymphulinae) were observed producing a high-intensity calling song (95 dB SPL at a distance of 10 cm) with a complex amplitude and frequency modulation (peaks of carrier frequency at 60 and 120 kHz). This sound is produced by a hitherto unknown type of sound organ located in the last abdominal (genital) segment, which may act as a tymbal. The observed directionality of sound output is probably achieved by means of a hollow cone surrounding the sound organ. Electrophysiological recordings revealed that the tympanal organs of S. minoralis are most sensitive in the frequency range from 50 to at least 100 kHz, which is distinctly higher than the minimum threshold levels in most other moths yet examined. The origin of genital sound production is discussed with respect to abdominal pheromone glands and pheromone-releasing movements.

1 aHeller, Klaus-Gerhard1 aKrahe, Rüdiger uhttp://jeb.biologists.org/content/jexbio/187/1/101.full.pdf01003nas a2200133 4500008004100000245008700041210006900128300001000197490000600207520054100213100002600754700002000780856006900800 1993 eng d00aThe ultrasonic song of the moth Amyna natalis (Lepidoptera:Noctudidae: Acontiinae)0 aultrasonic song of the moth Amyna natalis LepidopteraNoctudidae  a89-970 v53 a

In Malaysia, males of the noctuid moth Amyna natalis were observed producing a continuous ultrasonic song of high intensity (about 102 dB SPL measured at a distance of 10cm). The frequency spectrum of the sound impulses had its peak between 60 and 80 kHz. During song production the animals were perching on plants and moving their wings up and down quickly. Simultaneously, by twisting the wings it seems likely that a male-specific “bubble'' in the forewing functions as a tymbal, resulting in sound production.

1 aHeller, Klaus-Gerhard1 aAchmann, Roland uhttp://www.tandfonline.com/doi/abs/10.1080/09524622.1993.975323102304nas a2200121 4500008004100000245012100041210006900162300001200231490000700243520167100250100002601921856023501947 1996 eng d00aEcholocation and body size in insectivorous bats: the case of the giant naked bat Cheiromeles torquatus (Molossidae)0 aEcholocation and body size in insectivorous bats the case of the a27–380 v113 a

Recent hypotheses propose that the maximum size of aerial insectivorous bats is limited by properties of their echolocation system. According to these hypotheses large bats would (A) have to use low call frequencies which may only allow the detection of large prey. However, large prey may be too scarce to meet a large bat’s energy requirements. Large bats would (B) have low wing beat frequencies and due to energetic reasons also low call repetition rates. These low call repetition rates, however, many not be sufficient for prey detection. Both hypotheses were tested by analysing the echolocation behaviour of Cheiromeles torquatus (150-200g), the largest aerial insectivorous bat.

Cheriomeles searches for prey by alternating calls with the lowest frequencies at 20 and 28 kHz (call duration 30 and 24 ms respectively) at a repetition rate of 6.3Hz, and a duty cycle of 18%. The data are discussed in relation to the foraging behaviour of Cheiromeles.

The call parameters of Cheiromeles are far removed from predictions made in both models When the data of Cheiromeles and some other large bats are included in an allometric version of model (A) the prediction of the model changes distinctly indicating that call frequency might only be a weak constraint on body size, if at all. Call repetition rate seems to be correlated to call frequency and not directly to body size as is assumed in model (B). In nay case, large bats have lower call repetition rates than wing beat rates and the difference increases with body size. Therefore other factors have to be sought to explain the size distribution of aerial insectivorous bats.

1 aHeller, Klaus-Gerhard uhttps://www.researchgate.net/profile/Klaus-Gerhard_Heller/publication/257655672_Echolocation_and_body_size_in_insectivorous_bats_the_case_of_the_giant_naked_bat_Cheiromeles_torquatus_(Molossidae)/links/0deec52b5d412785f3000000.pdf00322nas a2200121 4500008004100000022002200041245001700063210001700080260002500097300001100122100002600133856004100159 2016 eng d a978-2-9537533-9-400aBioacoustics0 aBioacoustics aCannesbEdition GEEM a77-1001 aHeller, Klaus-Gerhard uhttps://bio.acousti.ca/da/node/1518301885nas a2200121 4500008004100000245010200041210006900143300001100212490000800223520146500231100002601696856004101722 1984 eng d00aZur Bioakustik und Phylogenie der Gattung Poecilimon (Orthoptera, Tettigoniidae, Phaneropterinae)0 aZur Bioakustik und Phylogenie der Gattung Poecilimon Orthoptera  a69-1170 v1113 a

35 European bush cricket species of the genus Poecilimon are comapred with special reference to their stridulatory movements, recorded with an opto-electronic camera, analysed in oszillograms and some power spectra, and ot their stridulatory files, studied with an electrone microscope. Since in the bush crickets the acoustics play the prime role in pair finding and thus reproductive isolation, an anslysis based on sound production and additional morphological characetristics lends itself well to investigate the relationships and the status of the species within this group. From this certain reclassifications are proposed: 15 species have been synonymized with 7 of them classified as supspecies and one subspecies has been newly described as Poecilimon obesus artedentatus.

The evolution of striduylatory movements and songs id sicussed extensively (Fig. 27). It is implied that in the songs shortneing or lengthening or fast repition of elements are typical derived forms from the original pattern. The phylogenetic tree presented in Fig. 26 considers differences in teh above physiological characteristcs as well as certian morphological onesexpecially the expression of the prothoracal stigma, whether wide or of a slit shape, the form of the pronotum, whether flat or raised, the form of the cerci, whether smooth or toothed, and finally it considers also the characteristic form of the animals stridulatory file.

1 aHeller, Klaus-Gerhard uhttps://bio.acousti.ca/da/node/15116