02065nas a2200301   4500008004100000022001400041245012200055210006900177260001600246300000800262490000900270520095900279653001301238653001001251653001801261653002601279653002701305653003501332653001601367100002101383700002801404700002801432700002001460700002101480700001601501700001901517856022701536       2017                        eng d  a1175-532600aYet another small brown frog from high altitude on the Marojejy Massif, northeastern Madagascar (Anura: Mantellidae)0 aYet another small brown frog from high altitude on the Marojejy   cFeb-11-2018  a5720 v43473 aMadagascar hosts a high diversity of small brown frogs. In this paper, we add another one by describing Gephyromantis (Duboimantis) tohatra sp. nov. The new species is a small brown mantellid frog discovered on a recent expedition to Marojejy National Park in northeastern Madagascar. It is characterised, among other things, by its small size (snout-vent length ~33 mm), an orange to yellowish belly, two dorsolateral ridges, and a distinctive call composed of 7–10 pulsed notes. The new species occurs sympatrically with other members of the subgenus Duboimantis at high altitude (~1700 m above sea level), including its sister species G. schilfi from which it radically differs by advertisement call and by a substantial genetic divergence of 4.3% uncorrected pairwise distance in the mitochondrial 16S rRNA gene. It thus joins the diverse assemblage of Gephyromantis species known from high altitudes on the mountain massifs of northern Madagascar.10aAmphibia10aAnura10aGephyromantis10aGephyromantis schilfi10aGephyromantis tandroka10aGephyromantis tohatra sp. nov.10aMantellidae1 aScherz, Mark, D.1 aRazafindraibe, Jary, H.1 aRakotoarison, Andolalao1 aDixit, Nadi, M.1 aBletz, Molly, C.1 aGlaw, Frank1 aVences, Miguel  uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4347.3https://biotaxa.org/Zootaxa/article/view/zootaxa.4347.3.9https://biotaxa.org/Zootaxa/article/viewFile/34380/30388https://biotaxa.org/Zootaxa/article/viewFile/34380/3038802264nas a2200241   4500008004100000022001400041245010400055210006900159260001600228300000800244490000900252520147400261653001301735653002901748653001501777653002301792653002401815653001301839100001901852700001901871700001601890856011601906       2017                        eng d  a1175-532600aA new species of smooth-skinned Spinomantis frog (Anura: Mantellidae) from south-eastern Madagascar0 anew species of smoothskinned Spinomantis frog Anura Mantellidae   cApr-09-2017  a3790 v43173 a<p>We present molecular evidence for the presence of two species morphologically similar to Spinomantis bertini in Andohahela National Park, south-eastern Madagascar, differing by 5.5&minus;6.3% pairwise DNA sequence divergences in the mitochondrial 16S rRNA gene. One of these was observed at higher elevations of ca. 1650 m above sea level, whereas the other was found at lower elevations of ca. 715 m a.s.l., close to the type locality of S. bertini (Isaka-Ivondro), and in one other location (Andreoky, ca. 1050 a.s.l.). We herein assign these low- to mid-elevation specimens to S. bertini based on their occurrence near the type locality and general agreement in colour pattern with the type specimen of Gephyromantis bertini Guib&eacute;, 1947. The high-elevation form is described as Spinomantis beckei sp. nov. based on its molecular divergence and reciprocal monophyly with respect to S. bertini, lower expression of greenish dorsal colour and less distinct frenal stripe. Based on a comparison of published call descriptions for S. bertini and our recordings of S. beckei, we hypothesize that S. bertini has a lower note repetition rate in advertisement calls. Molecular data suggest that the S. bertini species complex is more diverse than previously recognized, with at least two more candidate species identified: S. sp. Ca7 from Ranomafana National Park, and a newly identified candidate species S. sp. Ca12 from Pic d&rsquo;Ivohibe Special Reserve.</p>
10aAmphibia10aAndohahela National Park10aMadagascar10aSpinomantis beckei10aSpinomantis bertini10ataxonomy1 aVences, Miguel1 aKöhler, Jörn1 aGlaw, Frank  uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4317.2https://biotaxa.org/Zootaxa/article/view/zootaxa.4317.2.12