Kerman K, Roggero A, Piccini I, Rolando A, Palestrini C. Dung beetle distress signals may be correlated with sex and male morph: a case study on Copris lunaris               (Coleoptera: Scarabaeidae, Coprini).  2020. Available at: https://www.tandfonline.com/doi/full/10.1080/09524622.2019.1710255https://www.tandfonline.com/doi/pdf/10.1080/09524622.2019.1710255.